Abstract: Madagocerum gen. n. is described from Madagascar, representing the second genus of Odontoceridae from the Afrotropical Biogeographical Region. The following new species are described: <I>M. bhemi</I>, <I>M. idvigum</I> and <I>M. thoderirk</I> spp. n. All species were collected from moderately high elevations (above 1,550 meters).
Abstract: The following new replacement names for homonyms in the genus Ecnomus MCLACHLAN are proposed: Ecnomus tugelensis nom. nov. for Ecnomus dentatus Jacquemart, 1963; Ecnomus jacquemarti nom. nov. for Ecnomus ulmeri (JACQUEMART, 1961).
Abstract: Background
Leptoceridae are among the three largest families of Trichoptera (caddisflies). The current classification is founded on a phylogenetic work from the 1980's, based on morphological characters from adult males, i.e. wing venation, tibial spur formula and genital morphology. In order to get a new opinion about the relationships within the family, we undertook a molecular study of the family based on sequences from five genes, mitochondrial COI and the four nuclear genes CAD, EF-1alpha, IDH and POL.
Results
The resulting phylogenetic hypotheses are more or less congruent with the morphologically based classification, with most genera and tribes recovered as monophyletic, but with some major differences. For monophyly of the two subfamilies Triplectidinae and Leptocerinae, one tribe of each was removed and elevated to subfamily status; however monophyly of some genera and tribes is in question. All clades except Leptocerinae were stable across different analysis methods.
Conclusions
We elevate the tribes Grumichellini and Leptorussini to subfamily status, Grumichellinae and Leptorussinae, respectively. We also propose the synonymies of Ptochoecetis with Oecetis and Condocerus with Hudsonema.
Abstract: Male and female of Goera pitisopai sp. nov. from the Solomon Islands are illustrated and described based on recently collected material. This is the first species of the family Goeridae reported from the Solomon Islands, and the sixth from the Australasian region.
Abstract: The Trichoptera diversity of Vanuatu is reviewed based on earlier and recently collected
material. Altogether, 20 species in 12 genera and 8 families are now recognized
for the country, of which 12 species are described as new to science in the families
Hydrobiosidae (1 species), Hydroptilidae (5 species), Philopotamidae (3 species) and
Leptoceridae (3 species). The previously known species from Vanuatu are re-described
with illustrations and photographs. A key to adults of the Trichoptera fauna of Vanuatu
is provided.
Abstract: The genus <I>Apsilochorema</I> Ulmer, 1907 is unique in the family Hydrobiosidae Ulmer, being widely distributed in the Palaearctic, Oriental and Australian Regions. All other 49 genera in the family, except the New World <I>Atopsyche</I> Banks, 1905, are confined to a single biogeographical Region. This unique distribution has independently stimulated researchers to formulate competing hypotheses about the biogeographical history of the genus. Molecular sequence data from mitochondrial cytochrome oxidase I (COI) and nuclear cadherin (CAD) genes of <I>Apsilochorema</I> species from the Oriental and Australian areas were analysed phylogenetically. The results retain a monophyletic <I>Apsilochorema</I>, which forms the sistergroup to the other genera in the subfamily Apsilochorematinae. The results from the biogeographical analyses dispute the earlier assumptions of an Oriental or northern Gondwana origin for the genus, revealing unambiguously an initial Australian radiation of the ancestral <I>Apsilochorema</I> with a subsequent dispersal into the Oriental Region. All but one of the <I>Apsilochorema</I> species occurring on the Pacific islands had an Oriental ancestor. The exception is the sistergroup to the New Caledonian species, which is found in both Australia and Oriental Regions. The molecular dating analysis, using a relaxed clock model, indicates that the genus <I>Apsilochorema</I> is about 36.4 MY old and that it dispersed from Australia into the Oriental Region about 28.3 Ma. It also gives an estimate of the approximate ages of the dispersals into New Caledonia to about 15.3 Ma; to the Solomon Islands at about 16.2 Ma; to the Fiji Islands at about 16.1 Ma; and to the Vanuatu Islands at about 5.4 Ma.
Abstract: A new ecnomid genus and species, <I>Caledomina noumea</I> n. g. et n. sp. is described. The establishment of the new genus is based on its phylogenetic position in relation to other taxa in the family, but is also distinguished from the other genera in the family by several morphological character states in the males. A key to the genera in the family is provided based on males. The phylogenetic
position of the new genus is indicated based on previously published analyses of molecular data.
Abstract: The following new species are described from the west coast of Sumatra (Indonesia): Dipseudopsis gabiana and Pseudoneureclipsis complexa (Dipseudopsidae); Psychomyia selatana and Tinodes kerinciana (Psychomyiidae); Nyctiophylax anryutah, N. kupirah, and N. tapan (Polycentropodidae); Diplectrona dimagaha, D. tiguna, Hydropsyche pulakita, and Cheumatopsyche urhpirdana (Hydropsychidae); and Oecetis barisan, Setodes pantangulil, S. trikuthia, and S. wirhal (Leptoceridae). In addition, the following species are recorded from Sumatra for the first time: Chimarra briseis Malicky (Philopotamidae); Ecnomus hendersoni Mosely (Ecnomidae); Diplectrona dulitensis Kimmins and Hydropsyche butes Malicky & Chantaramongkol (Hydropsychidae); and Ganonema ochraceellum (McLachlan) (Calamoceratidae). Fourteen more species are recorded, but are previously known from Sumatra.
Abstract: Forty-six new species in the hydroptilid genera Acroptila WELLS, 1982 (2 spe- cies), Hellyethira NEBOISS, 1977 (2), Hydroptila DALMAN, 1819 (5), Microptila RIS, 1897 (1), Oxyethira EATON, 1873 (4), Caledonotrichia SYKORA, 1967 (1), Orthotrichia EATON, 1873 (14), Catoxyethira ULMER, 1912 (1), Chrysotrichia SCHMID, 1958 (8), Plethus HAGEN, 1887 (2), Scelotrichia ULMER, 1951 (3), Stactobia MCLACHLAN, 1880 (2), and Stactobiella MARTYNOV, 1924 (1) are described and illustrated for the first time. The ge- nus Parastactobia SCHMID, 1958 is synonymised with Catoxyethira ULMER, 1912. Hydro- ptila triangula XUE et YANG, 1990 is synonymized with H. thaphena OLAÌH, 1989. Hydroptila molione MALICKY, 2004 is synonymised with H. thuna OLAÌH, 1989. Two species, Oxy- ethira columba (NEBOISS, 1977) and Hydroptila keres MALICKY, 2004 are redescribed. New records for 52 species are given. With 197 figures.
Abstract: In area, New Caledonia is the smallest of the worldâs 25 official biodiversity hotspots, but in many taxonomic groups, the island has the highest concentration of species on earth, particularly so in the freshwater insect order Trichoptera. This study aims at applying molecular data and morphology for estimating the real species diversity of the genus Agmina on New Caledonia and investigating potential effects of ultramafic rock substrate on diversifica- tion. A dated molecular phylogeny was applied to study diversity and diversification related to geological substrate using the dispersalâextinctionâ cladogenesis model, D I V A and Bayesian ancestral character reconstruction. More than 47 species (> 63%) were unknown to science. Initial radiation occurred on ultramafic substrate followed by several independent dispersal events to nonultramafic substrate. The rate of shift from ultramafic to nonultramafic substrate was significantly higher than the rate of shift in the
opposite direction, ultramafic substrate.
indicating a possible cost associated with living on
Abstract: Nine new species of Chimarra (Philopotamidae) are described from the Solomon Islands. Chimarra maculata, C. veveensis, C. babarensis, and C. ventrospina are described from Kolombangara Island; and C. talinensis, C. kolombangensis, C. vitapinensis, C. solomonensis, and C. rosavensis were described from both Guadalcanal Island and Kolombangara Island. Chimarra biramosa Kimmins was collected from both Kolombangara Island and Guadalcanal Island.
Abstract: Nineteen new species of Dipseudopsidae from the Oriental Region are described and illustrated, including 6 new species of the genus Dipseudopsis: D. cocon, D. oliveri, D. gunung, D. rathnotoia, D. vienha, and D. viklundi; 2 new species of the genus Hyalopsyche: H. trunga and H. orissa; and 11 new species of the genus Pseudoneureclipsis: P. bonkybin, P. boquan, P. cauky, P. congkem, P. dongian, P. hailan, P. halongensis, P. loang, P. puyah, P. quancong, and P. vetcat. The genus Hyalopsychella Ulmer, 1930, is recognized as a new synonym of Hyalopsyche Ulmer, 1904. Hyalopsyche parvispinosa Schmid, 1959, and H. parvula Martynov, 1935, are recognized as new synonyms of H. sachalinica Martynov, 1910; Hyalopsyche similis Martynov, 1935, is recognized as a new synonym of Hyalopsychodes rivalis Betten, 1909; and Hyalopsychella haplotes Neboiss, 1989, is recognized as a new synonym of Hyalopsyche winkleri (Ulmer, 1930). New records of 14 other species are provided.
Abstract: Thirteen species of Helicopsyche, subgenus Feropsyche (Trichoptera: Helicopsychidae) are recorded from Costa Rica, including 3 new species: H. alajuela, n. sp., H. dorsocurvata, n. sp., and H. golfitoensis, n. sp., as well as 10 previously described species: H. borealis (Hagen), H. chiriquensis Johanson & Malm, H. dampfi Ross, H. incisa Ross, H. lewalleni Denning & Blickle, H. mexicana Banks, H. rentzi Denning & Blickle, H. selanderi Ross, H. truncata Ross, and H. vergelana Ross. The male genitalia of the new species are illustrated and distribution maps are provided for all species.
Abstract: Aim To test whether environmental diversification played a role in the diversification of the New Caledonian Hydropsychinae caddisflies.
Location New Caledonia, south-west Pacific.
Methods The phylogeny of the New Caledonian Hydropsychinae caddisflies was hypothesized using parsimony and Bayesian methods on molecular characters. The Bayesian analysis was the basis for a comparative analysis of the correlation between phylogeny and three environmental factors: geological substrate (ultrabasic, non-ultrabasic), elevation and precipitation. Phylogenetic divergence times were estimated using a relaxed clock method, and environmental factors were mapped onto a lineage-through-time plot to investigate the timing of environmental diversification in relation to species radiation. The correlation between rainfall and elevation was tested using independent contrasts, and the gamma statistic was calculated to infer the diversification pattern of the group.
Results The diversification of extant OrthopsycheâCaledopsyche species began in the MiddleâLate Oligocene, when much of the island of New Caledonia was covered by ultrabasic substrate and mountain forming was prevalent. Most lineages originated in the MiddleâLate Miocene, a period associated with long-term climate oscillation. Optimization of environmental factors on the phylogeny demonstrated that the New Caledonian Hydropsychinae group adapted to ultrabasic substrate early in its evolutionary history. The clade living mostly on ultrabasic substrate was far more species-rich than the clade living mostly on non-ultrabasic substrate. Elevation and rainfall were significantly correlated with each other. The lineage- through-time plot revealed that the main environmental diversification preceded species diversification. A constant speciation through time was rejected, and the negative gamma indicates that most of the diversification occurred early in the history of the clade. According to the inferred phylogeny, the genus Orthopsyche McFarlane is a synonym under Caledopsyche Kimmins, and Abacaria caledona Oláh & Barnard should also be included in Caledopsyche.
Main conclusions The age of the radiation does not support a vicariance origin of New Caledonian Hydropsychinae caddisflies. Environmental diversification pre-dates lineage diversification, and thus environmental heterogeneity potentially played a role in the diversification of the group, by providing a variety of fragmented habitats to disperse into, promoting speciation. The negative gamma indicates that the speciation rate slowed as niches started to fill.
Abstract: The taxonomy of the Polycentropodidae is discussed, and the family is divided into 4 newly defined diagnostic genus clusters based primarily on wing characters and number of spurs on the legs. The diagnostic genus clusters are the Neureclipsis diagnostic genus cluster with Neureclipsis McLachlan and Neucentropus Martynov; the Polycentropus diagnostic genus cluster with Polycentropus Curtis, Plectrocnemia Stephens, Holocentropus McLachlan, and Polyplectropus Ulmer; the Cyrnus diagnostic genus cluster with Nyctiophylax Brauer, Cyrnus Stephens, Cyrnopsis Martynov, Cyrnellus Banks, and Adectophylax Neboiss; and the Cyrnodes diagnostic genus cluster with Cyrnodes Ulmer, Cernotina Ross, and Pahamunaya Schmid. The extinct genus Archaeoneureclipsis Ulmer is synonymised with Neureclipsis McLachlan; the extant genus Tasmanoplegas Neboiss is synonymised with Plectrocnemia Stephens; the extant genus Eodipseudopsis Marlier is synonymised with Polyplectropus Ulmer; and the extinct genus Nyctiophylacodes Ulmer is synonymized with Nyctiophylax Brauer. The following 8 new species are described in Plectrocnemia: P. martynovi (Myanmar), P. ambaita (Myanmar), P. kachin (Myanmar), P. taungyia (Myanmar), P. dalat (Vietnam), P. kamba (Myanmar), P. malaisei (Myanmar), and P. thai (Vietnam). The following 11 new species are described in Polyplectropus: P. daimong (Vietnam), P. dinhdan (Vietnam), P. giandi (Vietnam), P. jalan (Malaysia), P. pairavatika (Malaysia), P. vanuatu (Vanuatu), P. coronivia (Fiji), P. palma (Fiji), P. vanda (Fiji), P. vanua (Fiji), and P. wainimbuk (Fiji). The following 2 new species are described in the subgenus Nyctiophylax (Nyctiophylax): N. (N.) hotay (Vietnam) and N. (N.) catunujah (Myanmar). The following 7 new species are described in the subgenus Nyctiophylax (Paranyctiophylax): N. (P.) anosib (Madagascar), N. (P.) buoc (Vietnam), N. (P.) dhauli (India), N. (P.) kilah (Madagascar), N. (P.) lancelot (Malaysia), N. (P.) mintin (Vietnam and Laos), N. (P.) nepenthes (Malaysia). Cyrnopsis tangaron is described from Indonesia (Borneo). And the following 3 new species in Pahamunaya are described: P. espelandae (Brunei), P. wamana (Malaysia) and P. khoii (Vietnam). In addition, new records for 20 species are given.
Abstract: Additions to the taxonomy of the leptoceroid families Calamoceratidae, Molannidae, Odontoceridae, and Philorheithri- dae are given, and 33 new species in the superfamily are described based on characters in the male genitalia, head and thoracal setose warts, and groove patterns, wing venation, and forewing colour and pattern.
In the Calamoceratidae Ulmer, the Anisocentropus latifasciata diagnostic species-group and Anisocentropus brevi- pennis diagnostic species cluster are described for the first time. The following 17 species are described as new: Anisoce- ntropus bungus (Vietnam), A. csorbai (Vietnam), A. dvaupadhah (Malaysia), A. fridae (Fiji Islands), A. hannahae (Fiji Islands), A. hoisat (Laos PDR), A. malaisei (Myanmar), A. maralus (Malaysia), A. mjoebergi (Borneo), A. samuh (Mad- agascar), A. tapenan (Solomon Islands), A. thinlin (Laos PDR), A. thonmihn (Vietnam), A. vanuensis (Fiji Islands), A. vitiensis (Fiji Islands), Ganonema malickyi (Thailand) and G. rong (Vietnam). In addition, Ganonema fuscipenne (Albarda) is synonymized with G. ochraceellum (McLachlan), and Ganonema pallidum Martynov is synonymized with Anisocentropus kawamurai (Iwata).
In the Molannidae Wallengren, a single new species, Molanna gamdaha, is described from Myanmar.
In the Odontoceridae Wallengren, the genus Inthanopsyche Malicky is synonymized with Psilotreta Banks. The fol- lowing 14species belonging to previously described genera, are described as new: Lannapsyche birathena (Myanmar), L. suksma (Myanmar), L. kamba (Myanmar), Marilia enikiana (Laos PDR), M. jonssoni (Laos PDR), M. katakaha (Malay- sia), M. malickyi (Laos PDR, Malaysia, Vietnam), M. mendolonga (Malaysia), M. namha (Laos PDR), M. tuyetmira (Laos PDR), Phraepsyche pectinata (Vietnam), P. yitungshana (China: Hong Kong), Psilotreta enikoae (Vietnam), and P. malickyi (Myanmar). In addition, the following 3 diagnostic species-groups are described for the first time: Psilotreta japonica diagnostic species-group, Psilotreta trimeresuri diagnostic species-group, and Psilotreta frontalis diagnostic species group. The species Ganonema odaenum Kobayshi is synonymized with Psilotreta locumtenens Botosaneanu, and Psilotreta pyonga OlaÌh is synonymized with P. falcula Botosaneanu.
In the Philotheithridae Mosely, a single new species, Psilopsyche granda, is described from Chile. In addition to describing new taxa, new species records for 34 species in the superfamily are presented.
Abstract: Six new species of Paduniella (Insecta, Trichoptera, Psychomyiidae) from Asia are described and illustrated: P. burmana (Burma), P. dongdata (Vietnam), P. monga (Vietnam), P. nama (Vietnam), P. sabarmata (India), and P. taya (Vietnam). The male genitalia of P. sampati Malicky & Chantaramongkol and P. semarangensis Ulmer are re-described and illustrated based on new material from Vietnam.
Abstract: Background: The insecticides dichlorvos, paradichlorobenzene and naphthalene have been commonly used to eradicate pest insects from natural history collections. However, it is not known how these chemicals affect the DNA of the specimens in the collections. We thus tested the effect of dichlorvos, paradichlorobenzene and naphthalene on DNA of insects (Musca domestica) by extracting and amplifying DNA from specimens exposed to insecticides in two different concentrations over increasing time intervals.
Results: The results clearly show that dichlorvos impedes both extraction and amplification of mitochondrial and nuclear DNA after relatively short time, whereas paradichlorobenzene and naphthalene do not.
Conclusion: Collections treated with paradichlorobenzene and naphthalene, are better preserved concerning DNA, than those treated with dichlorvos. Non toxic pest control methods should, however, be preferred due to physical damage of specimens and putative health risks by chemicals.
Abstract: Five new species of Paduniella (Trichoptera: Psychomyiidae) are described: P. flinti, P. nandra, P. ambra, P. madagassa, and P. sona, representing the first records of the genus from Madagascar and more than doubling the number of Paduniella species known from the Afrotropical Region.
Abstract: Calocidae constitute a hypothesised monophyletic group of caddisflies (Trichoptera) being geographically restricted to New Zealand (one genus) and Australia (five genera). This analysis tests the monophyly of the family based on sequences from five different molecular genes. The complete data set includes 29 species and covers a complete genus representation of the Calocidae as well as representatives of other families in which one or more calocid genera have been classified. Sequences from two mitochondrial (cytochrome oxidase I and 16S) and three nuclear (elongation factor 1-a, RNA polymerase-II, and Cadherin) genes were used, resulting in a 3958 bp data set and 37.1% parsimony informative characters. The Cadherin (CAD) and RNA polymerase-II (POL-II) genes are used for the first time for revealing Trichoptera phylogenies. The character matrix was analyzed by using maximum parsimony (MP) and Bayesian criteria, the latter by applying three different partition strategies for comparison. Two most parsimonious trees were found, differing in the position of one clade within the sister-group to a monophyletic Calocidae. The Bayesian tree based on the maximum number of partitions differs from trees based on a reduced partition analysis with respect to taxa outside the current circumscription of Calocidae. Both the MP and Bayesian analyses left Calocidae monophyletic, with a monophyletic clade of all Australian genera being sister-group to the New Zealand genus. The results from the agreement subtree analysis demonstrates that CAD performs well both separately and in combination with other genes and adds substantial resolution to the calocid phylogeny in a combined MP analysis.
Abstract: Ecnomidae are a family of seven previously accepted extant genera having a typical Gondwanan distribution, except one genus (Ecnomus) being widely distributed also in the Oriental and Palearctic regions. We analysed a molecular data set of 3379 characters representing the sum of four different protein-coding genes (COI, CAD, EF-1a and POL-II). Six equally most parsimonious trees were generated from the combined data set, distributed into two distinct islands. In all maximum parsimony (MP) trees the Ecnomidae is monophyletic when the genus Zelandoptila (Psychomyiidae) is included. The sister group to Ecnomidae including Zelandoptila is Pseudoneureclipsis, previously classified in the other families. This sister-group relationship contradicts earlier findings that the Polycentropodidae are the sister group to Ecnomidae. A Bayesian analysis resulted in a monophyletic Ecnomidae when accepting inclusion of Pseudoneureclipsis, which contradicts the results from the MP analysis by leaving Zelandoptila as the sister group to Ecnomidae including Pseudoneureclipsis. In the majority rule tree from this analysis Polycentropodidae form the sister group to Ecnomidae. We were not able to obtain a monophyletic Ecnomus due to the inclusion of Psychomyillodes. We conclude that the genus Zelandoptila or Pseudoneureclipsis probably belongs to the Ecnomidae, and that Psychomyiellodes and Ecnomus are synonyms. Three additional, as yet undescribed monotypic genera from Australia and New Caledonia remain to be erected in Ecnomidae.
Abstract: The following sixteen Trichoptera species new to science are described and illustrated: Cheumatopsyche navasi sp. n., C. olahi sp. n., C. barnardi sp. n., Potamyia schlingeri sp. n., Leptonema irwini sp. n. (Hydropsychidae Curtis), Nyctiophylax madagassa sp. n. (Polycentropodidae Ulmer), Chimarra apiconigra sp. n., C. blahniki sp. n., C. gassa sp. n., Wormaldia curvus sp. n., W. leviovatus sp. n., W. trispina sp. n. (Philopotamidae Stephens), Goera madagassa sp. n. (Goeridae Ulmer), Lepidostoma asymmetrica sp. n. (Lepidostomatidae Ulmer), Pisulia lata sp. n. and P. magna sp. n. (Pisuliidae Ross). The families Polycentropodidae and Goeridae are recorded from Madagascar for the first time. With this report, 204 Trichoptera species are now known from the island. A first key to adults of the known families and genera of Madagascan Trichoptera is provided.
Abstract: Twenty-one new Polycentropodidae (Trichoptera) species are described: Polyplectropus aberrus, P. dorsospinus, P. nodyg, P. yndog, P. clavus, P. nathalae, P. millei, P. christinae, P. koueus, P. viklundi, P. hovmoelleri, P. aoupiniensis, P. tenerus, P. angustus, P. curvispinus, P. caledonia, P. piroguensis, P. triangulatus, P. pernodensis, P. taoensis, and P. papei spp. novae, representing the first species records of this family from New Caledonia. A key to males of the New Caledonian Polycentropodidae is provided, and distribution maps are presented for all species.
Abstract: We report five new species records from the Comoros Archipelago. Two of the species are known from outside the Archipelago, Hydroptila cruciata Ulmer (Hydroptilidae) and Anisocentropus voeltzkowi Ulmer (Calamoceratidae), and three species are described as new to science: Pisulia stoltzei, new species (Pisuliidae), and: Chimarra mayottensis, new species and Chimarra koulaeensis, new species (Philopotamidae). Five species have been previously recorded from the Comoros Islands: Cheumatopsyche comorina (NavaÌs), Macrostemum capense (Walker), Cheumatopsyche vala Malicky (Hydropsychidae), Hydroptila voticia Malicky (Hydroptilidae), and Oecetis atpomarus Malicky (Leptoceridae). With this report the number of species from the Comoros is doubled. These findings also represent the first records of Trichoptera from Mayotte.
Abstract: Conoesucidae (Trichoptera, Insecta) are restricted to SE Australia, Tasmania and New Zea- land. The family includes 42 described species in 12 genera, and each genus is endemic to either New Zealand or Australia. Although monophyly has been previously assumed, no morphologi- cal characters have been proposed to represent synapomorphies for the group. We collected molecular data from two mitochondrial genes (16S and cytochrome oxidase I), one nuclear gene (elongation factor 1-a) (2237â2277 bp in total), and 12 morphological characters to pro- duce the first phylogeny of the family. We combined the molecular and morphological charac- ters and performed both a maximum parsimony analysis and a Bayesian analysis to test the monophyly of the family, and to hypothesize the phylogeny among its genera. The parsimony analysis revealed a single most parsimonious tree with Conoesucidae being a monophyletic taxon and sistergroup to the Calocidae. The Bayesian inference produced a distribution of trees, the consensus of which is supported with posterior probabilities of 100% for 15 out of 22 possible ingroup clades including the most basal branch of the family, indicating strong sup- port for a monophyletic Conoesucidae. The most parsimonious tree and the tree from the Bayesian analysis were identical except that the ingroup genus Pycnocentria changed position by jumping to a neighbouring clade. Based on the assumption that the ancestral conoesucid spe- cies was present on both New Zealand and Australia, a biogeographical analysis using the dis- persal-vicariance criteria demonstrated that one or two (depending on which of the two phylogenetic reconstructions were applied) sympatric speciation events took place on New Zealand prior to a single, late dispersal from New Zealand to Australia.
Abstract: Based on a collection of Trichoptera from the Finaransoa Province on Madagas- car, the following three new species are described and illustrated: Pseudoneureclip- sis madagascariensis, spec. nov. (Dipseudopsidae), Madagocerum flinti, spec. nov. (Odontoceridae), and Cheimacheramus ranomafanensis, spec. nov. (Sericostomati- dae).
This report represents the first record of the genus Pseudoneureclipsis Ulmer from Madagascar, the fourth known species of the endemic Madagascan genus Madagocerum OlaÌh & Johanson, and the third known species in the Madagascan and South African genus Cheimacheramus Barnard.
Abstract: The taxonomy of the genera of Hydropsychinae is revised. The following genera are synonymized: Hydatomanicus Ulmer, 1951 and Hydatopsyche Ulmer, 1926 are synonymized with Hydromanicus Brauer, 1865; Streptopsyche Ross & Unzicker, 1977 is synonymized with Calosopsyche Ross & Unzicker, 1977; Symphitopsyche Ulmer, 1907, Mexipsyche Ross & Unzicker, 1977, Ceratopsyche Ross & Unzicker, 1977 and Herbertorossia Ulmer, 1957 are synonymized with Hydropsyche Pictet, 1834; and Plectropsyche Ross, 1947 is synonymized with Cheumatopsyche Wallengren, 1891. A new genus, Schmidopsyche OlaÌh & Schefter, new genus is established and the single species in the genus, S. rossi OlaÌh & Schefter, new species is described and illustrated, based on material collected by Schmid in Sikkim in 1959. The additional 61 new species are described and illustrated: India: Hydromanicus krsamgin OlaÌh & Barnard, new species; H. sacso OlaÌh & Johanson, new species; H. naraik OlaÌh & Barnard, new species; H. palnis OlaÌh & Barnard, new species; H. topali OlaÌh & Johanson, new species; Hydropsyche januha OlaÌh & Barnard, new species; H. kamenga OlaÌh & Schefter, new species; H. mizora OlaÌh & Schefter, new species; H. narayana OlaÌh & Johanson, new species; H. khasigiri OlaÌh & Barnard, new species; H. igunapali OlaÌh & Schefter, new species; H. kangra OlaÌh & Barnard, new species; H. kiogupa OlaÌh & Schefter, new species; H. minutanga OlaÌh & Johanson, new species; H. tritiyaha OlaÌh & Barnard, new species; H. ekaropa OlaÌh & Schefter, new species; H. yathetima OlaÌh & Schefter, new species; H. keralana OlaÌh & Barnard, new species; H. semkala OlaÌh & Johanson, new species; H. shillonga OlaÌh & Barnard, new species; Cheumatopsyche barakambra OlaÌh & Johanson, new species; C. mariannae OlaÌh & Johanson, new species; C. sandrae OlaÌh & Johanson, new species; Potamyia assamana OlaÌh & Schefter, new species. Vietnam: Hydromanicus demden OlaÌh & Johanson, new species; and Hydropsyche soinha OlaÌh & Johanson, new species (also from Malaysia); H. tanung OlaÌh & Johanson, new species; H. lomdom OlaÌh & Johanson, new species; H. sangbung OlaÌh & Johanson, new species; H. keoda OlaÌh & Johanson, new species; H. xenga OlaÌh & Johanson, new species; and Cheumatopsyche simayorum OlaÌh & Johanson, new species. Malaysia: Hydropsyche gekilara OlaÌh & Barnard, new species; H. tanua OlaÌh & Barnard, new species; and H. namea OlaÌh & Barnard, new species. Philippines: Hydropsyche meyi OlaÌh & Johanson, new species; Cheumatopsyche anonima OlaÌh & Johanson, new species; C. kuranishii OlaÌh & Johanson, new species; and C. montapo OlaÌh & Johanson, new species. Papua New Guinea: Hydropsyche flintorum OlaÌh & Johanson, new species; and H. carolae OlaÌh & Johanson, new species. Tibet: Hydropsyche gyantsana OlaÌh & Barnard, new species. Nepal: Hydropsyche nepalarawa OlaÌh & Johanson, new species; and Cheumatopsyche chitawana OlaÌh & Johanson, new species. China: Hydropsyche chengdu OlaÌh & Barnard, new species; and Cheumatopsyche yangmorseorum OlaÌh & Johanson, new species. Iran: Hydropsyche masula OlaÌh & Johanson, new species. New Caledonia: Orthopsyche anulmika OlaÌh & Johanson, new species. Japan: Cheumatopsyche okinawana OlaÌh & Johanson, new species; and C. tanidai OlaÌh & Johanson, new species. Taiwan: Cheumatopsyche davisi OlaÌh & Johanson, new species; and C. tungpa OlaÌh & Johanson, new species. Sri Lanka: Cheumatopsyche hasalaka OlaÌh & Johanson, new species. South Africa: Cheumatopsyche krugerana OlaÌh & Johanson, new species. Madagascar: Cheumatopsyche mafana OlaÌh & Johanson, new species; C. nondra OlaÌh & Johanson, new species; C. tiokalamkita OlaÌh & Johanson, new species; C. fianara OlaÌh & Johanson, new species; C. masiposa OlaÌh & Johanson, new species; C. ranoma OlaÌh & Johanson, new species; and C. rantsoa OlaÌh & Johanson, new species. The following species are synonymized: Hydropsyche kawamurai Tsuda, 1940 is synonymized with H. kozhantschikovi Martynov, 1924, Hydropsyche belos Malicky & Chantaramongkol, 2000 is synonymized with H. klimai Mey, 1999, Hydropsyche taiwanensis Mey, 1997 is synonymized with H. spinata Kobayashi, 1987, and Cheumatopsyche copiosa Kimmins, 1956 is synonymized with C. columnata Martynov, 1935. Recognized genera are divided into previously established and new species groups, defined by shared diagnostic characters. All Oriental species of Hydropsyche Pictet and Hydromanicus Brauer are grouped into different levels of species groups based on similar morphology in male genitalia.
Abstract: The toxic and nutrient poor ultrabasic rock substrate covering one-third of New Caledonia greatly influ- enced on the biogeography and diversity of plants in the island. Studies on the effect of ultrabasic sub- strate on fauna are almost entirely absent. In this paper we examine whether the diversification of Trichoptera of the New Caledonian endemic genus Xanthochorema Kimmins, 1953 was related to the presence of ultrabasic substrate. The analysis is based on data from a phylogeny derived from DNA sequences of mitochondrial COX1, COX2 and 16S, and nuclear EF1a genes. The study of the relationships between ancestral species and substrate was carried out using dispersalâvicariance analysis and tracing the history of substrate association with ultrabasic and non-ultrabasic distributions representing the ter- minals in the fully resolved phylogenetic tree. Our results show that (1) the ancestor of all Xanthochorema species was present on ultrabasic substrate, (2) early speciation events were restricted to ultrabasic sub- strate, (3) younger ancestral species dispersed into non-ultrabasic substrates, and (4) late speciation events were restricted to non-ultrabasic substrate. These results correspond to the hypothesis that New Caledonia once was more extensively covered by ultrabasic rocks than at present.
Abstract: A phylogenetic analysis is presented for Helicophidae genera based on fragments of the three genes COI, 16S and EF-1a (total 1705 bp) and morphology (70 characters). Molecular data show that both the family Helicophidae + Alloecentrella Wise and the New Caledonian species form monophyletic groups. Monophyly of the New Caledonian Helicophidae based on morphology alone is not supported. Nine new species of Helicopha (Trichoptera, Helicophidae) from New Caledonia are described: H. neocaledonia sp.n., H. loripes sp.n., H. angulata sp.n., H. stellata sp.n., H. bifurcata sp.n., H. mouirangensis sp.n., H. acuta sp.n., H. taoensis sp.n., and H. aoupiniensis sp.n. The Helicophidae now include 42 known species of which 18 are New Caledonian endemics. New Caledonian Helicophidae form a monophyletic group, whose species diversity is the greatest amongst the worldâs Helicophidae. Descriptions of the females of Helicopha dognyensis and H. ramea, previously known only from males, are the first for females from New Caledonia. Briama Johanson & Ward (syn.n.) is phylogenetically most closely related to New Caledonian Helicopha and is considered as synonymous. Alloecen- trella Wise from New Zealand, previously classified within Calocidae, is more closely related to Austrocentrus Schmid from Chile and is to be classified as Helicophidae.
Abstract: Helicopsyche agnetae, new species, is described and figured based on material collected in Hong Kong in May 2005. With this record two species of the family Helicopsychidae are known from China. The two Chinese species are probably sister species sharing several uniquely derived synapomorphies, and are separated from each other by features in the genitalia.
Abstract: Seven new Tinodes species are described from South-East Asia. Tinodes kemnounga, new species, Tinodes caolana, new species, Tinodes gapbona, new species, and Tinodes dactringa, new species are described from Vietnam. Tinodes samkuca, new species, Tinodes kebala, new species, and Tinodes suksa, new species are described from Peninsular Malaysia.
Abstract: The Oceanian country of New Caledonia has been shown to have a great diversity of Trichoptera, but prior to this work only 3 species from the large Leptoceridae genus Triplectides were known from there. Extensive sampling on the main island, Grande Terre, revealed 11 new species in the genus, as well as a male of the seemingly widespread species Triplectides australis. These 11 new species are here described and illustrated, and a key to males of the New Caledonian Triplectides species is provided. The new species are: T. mouiensis, new species; T. abnormalis, new species; T. minutus, new species; T. noumeiensis, new species; T. tigrinus, new species; T. koghiensis, new species; T. wardi, new species; T. nathaliae, new species; T. mariannae, new species; T. dawnae, new species; T. aequalichelatus, new species. Fifteen species within the genus are now known from New Caledonia; relative to land mass, this is a high diversity compared to the 25 species recorded from Australia.
Abstract: Three new species, Helicopsyche pathoumthongi, Helicopsyche vongsombathi, and Helicopsyche laoensis, are described and illustrated based on material collected in northern Laos from AprilâMay, 2005. With Helicopsyche antikleia Malicky, 1997, originally described from Laos, and the discovery of a single male of the widespread Oriental species Helicopsyche angusta (Ulmer, 1951) the Laotian Helicopsyche fauna now contains 5 recorded species.
Abstract: A method for associating larvae, females and males of Trichoptera is demonstrated for New Caledonian Hydrobiosidae species of the genus Xanthochorema, using cytochrome oxidase subunit I (COI) and 16S mitochondrial gene sequences. Two species, X. caledon Kimmins, 1953 and X. celadon Schmid, 1989, previously with unknown larvae and undescribed females, were associated, and males, females and larvae of both species are described. Mitochondrial COI and 16S gene fragments are demonstrated to be useful for association of sexes and life stages of the two species, and distance measures show that the method is likely to also be useful for other species within the genus. The associations are well supported by high bootstrap and jackknife values.
Abstract: Fourteen new Tinodes species are described from East Africa. Tinodes guttatus sp. n., Tinodes rungweensis sp. n., Tinodes sanjea sp. n., Tinodes amani sp. n., Tinodes robustus sp. n., Tinodes zigzag sp. n., Tinodes uzungwa sp. n. and Tinodes bumbulii sp. n. are described from the Eastern Arc mountains in Tanzania and represent the first described species of this genus from Tanzania. Tinodes harawa sp. n., Tinodes dehadawate sp. n., Tinodes mohelia sp. n., Tinodes irwini, sp. n., and Tinodes schlingeri sp. n. are described from Madagascar and the Comoros Island, and represent the first described species of this genus from these islands. Tinodes giloensis sp. n. is described from Sudan, and is the first record of Tinodes from the country. With this report the number of known Tinodes species from the Afrotropical region is more than doubled, and the Eastern Arc is made the richest area in number of species of Tinodes in that biogeographical region.
Abstract: Seven new species of Helicopsyche (Feropsyche) Johanson 2002 (Helicopsychidae) are described from Mexico (H. curvipalpia new species), Panama (H. blantoni new species, H. chiriquensis new species, H. linguata new species, and H. sanblasensis new species), and Brazil (H. paprockii new species and H. cipoensis new species) based on adult material borrowed from the National Museum of Natural History (Smithsonian Institution), Washington, D. C. and the Illinois Natural History Survey, Champaign, Illinois, USA. New records are given for H. sinuata Denning & Blickle from Mexico, and H. incisa Ross and H. woldai Johanson from Panama.
Abstract: Eleven new Trichoptera species of the superfamily Sericostomatoidea are described from Madagascar. Three species within Helicopsychidae are described as Helicopsyche (Petrotrichia) ambodiva, new species, Helicopsyche (Petrotrichia) sahadika, new species, and Helicopsyche (Petrotrichia) ninakosha, new species; six species within the Petrothrincidae are described as Petrothrincus tsaratananensis, new species, Petrothrincus newidop, new species, Petrothrincus dhritaparam, new species, Petrothrincus pauliani, new species, Petrothrincus andohel, new species, and Petrothrincus andring, new species; and two new species within the Sericostomatidae are described as Cheimacheramus rossi, new species, and Rhoizema mahalevonum, new species. The genus Gyrocarisa Weaver 1997 is synonymized with Petrothrincus Barnard 1934, resulting in the new combinations Petrothrincus steineri (Weaver 1997), new combination, Petrothrincus concava (Weaver 1997), new combination, Petrothrincus acuta (Weaver 1997), new combination, Petrothrincus scottae (Malm & Johanson 2005), new combination, and Petrothrincus weaveri (Malm & Johanson 2005), new combination. A key is presented to males of all Sericostomatoidea species from Madagascar. Distribution maps are given for all new and previously described Madagascar species within the superfamily.
Abstract: Three new species of Symphitoneuria Ulmer (Trichoptera: Leptoceridae) from New Caledonia are described and illustrated. These are Symphitoneuria lacsiensis, new species, Symphitoneuria digitata, new species and Symphitoneuria triangulata, new species. Together with these the genus now holds 11 species, and the New Caledonian Symphitoneuria now consists of 5 species, including the previously described Symphitoneuria licmetica Neboiss and Symphitoneuria clara Ward. A key to the males of the New Caledonian Symphitoneuria species is provided.
Abstract: The philopotamid genus Hydrobiosella is exclusively South-Pacific in distribution and restricted to Australia (20 species), New Zealand (4 species) and New Caledonia (6 species). The first species known from New Caledonia, Hydrobiosella uncinata Kimmins, 1953, was described from a single male collected in PueÌbo in 1949. In a recently collected material from New Caledonia H. uncinata together with five undescribed species were present. The genitalia of the H. uncinata holotype are re-described and the new species are described. Distribution maps and a key to the males are presented for all New Caledonian species.
